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Dry Oak-Hickory-Hophornbeam Forest

Ecology and Physical Setting illustration of dry oak hickory hophornbeam forest

These are open, park-like forests where shrubs are sparse—one can see a long distance through the woods. Such open forests are uncommon in Vermont, and visitors find them exceptionally beautiful. Dry Oak-Hickory-Hophornbeam Forests are found on hilltops, gentle ridgelines, knobs, benches, and other small rises. A combination of warm climates, shallow soils, and landforms that shed water creates a specific set of site conditions. Bedrock is various, ranging from calcareous to acidic, but there are almost always some plants indicating soil enrichment. Trees do not grow as well as they do on deeper, moister soils. Many forests of this type are found on the rocky portions of Champlain Valley farms. In most cases they were never cleared for pasture or crops but were often used as woodland pasture. Natural and anthropogenic disturbances probably contribute to the structure of Dry Oak-Hickory-Hophornbeam Forests. 

Vegetation 

Red oak, shagbark hickory, and hophornbeam are the common canopy dominants. Sugar maple, white ash, and white oak can be present, and even abundant, in the most enriched examples. The canopy is somewhat open, and trees do not reach great height. The shrub layer is very sparse. The ground layer is often a dense lawn of woodland sedge, with a diverse array of herbs scattered about. In areas of calcareous bedrock, rare species abound. 

Wildlife Habitatdry oak hickory hophornbeam forest

The melodic, flute-like call of the wood thrush and the plaintive call of the wood pewee are just some of the attractions of Dry Oak-Hickory-Hophornbeam Forests. Other nesting birds include white-breasted nuthatch, scarlet tanager, red-eyed vireo, rose-breasted grosbeak, and Baltimore oriole. The nutlets of hophornbeam are important food for turkeys, rose-breasted grosbeaks, gray squirrels, and chipmunks. Hickory nuts and the acorns of red and white oak are important for turkey, black bear, squirrels, and chipmunks. 

The entirely terrestrial eastern red-backed salamander may be abundant, especially in older forests with lots of downed and decaying wood. 

Three rare reptiles—timber rattlesnake, eastern ratsnake, and the five-lined skink—are all found in Dry Oak-Hickory-Hophornbeam Forests in specific areas of Vermont. Hickory is the primary host plant for the caterpillar stage of the uncommon hickory hairstreak butterfly. The large plates of exfoliating bark on mature shagbark hickory provide roosting sites for two rare bats: Indiana bat and northern long-eared bat.

Successional Trends

Sugar maple, red oak, white oak, shagbark hickory, and hophornbeam are all likely late-successional canopy components. Fire has played a role in some examples of this community, but whether fire is necessary to maintain these forests is not known.

Variants

  • Sugar Maple-Hophornbeam Forest also occurs on water-shedding ridgetops, but is found in slightly cooler and moister climates. It is dominated by sugar maple, with hophornbeam and red oak, but lacks shagbark hickory. White ash can be locally abundant. The structure and composition of the shrub and herb layers is very similar to typical Dry Oak-Hickory-Hophornbeam Forests.

Related Communities

  • Dry Oak Forest is more nutrient-poor or has bedrock closer to the surface, or both. Red oak, white oak, and chestnut oak dominate, and heath shrubs are common in the ground layer. Woodland sedge may be common in these forests, as it is in Dry Oak-Hickory-Hophornbeam Forest. 

     
  • Mesic Maple-Ash-Hickory-Oak Forest is found on lower slopes, where soils are deeper and moisture is more available to plants. The two communities often grade into each other. 

Conservation Status and Management Considerations

Dry Oak-Hickory-Hophornbeam Forest is an uncommon community, with only a handful of large, natural examples known. Smaller patches of this community type are moderately common in the Champlain Valley. Some excellent examples are protected on conserved lands. Calcareous examples of this community are especially vulnerable to invasion by non-native species. Any management activities should include efforts to control invasives.

Distribution/Abundancemap of Vermont with locations of natural community

This community has southern affinities and is found in the warmer parts of Vermont as well as in surrounding states. Most examples are in the Champlain Valley and Taconic Mountains, although the community is also known from the Southern Vermont Piedmont, and the Vermont Valley. Isolated occurrences may be found in some of the warmest areas of the Southern Green Mountains and the Champlain Hills.   

Characteristic Plants

Trees

Abundant Species 

Red oak – Quercus rubra

Hophornbeam – Ostrya virginiana

Shagbark hickory – Carya ovata

Occasional to Locally Abundant Species

White oak – Quercus alba

Sugar maple – Acer saccharum

White ash – Fraxinus americana

Shrubs

Occasional to Locally Abundant Species

Maple-leaved viburnum – Viburnum acerifolium

Downy arrowwood – Viburnum rafinesquianum

Herbs

four-leaved milkweed

Four-leaved milkweed.

Abundant Species

Woodland sedge – Carex pensylvanica

Occasional to Locally Abundant Species

Hog peanut – Amphicarpaea bracteata

Bottlebrush grass – Elymus hysterix

Blue-stemmed goldenrod – Solidago caesia

Pedunculate sedge – Carex pedunculata

Broad-leaved sedge – Carex platyphylla

Wild oats – Uvularia sessilifolia

Furry brome – Bromus pubescens

Blunt fescue - Festuca obtusa

Black-seeded mountain rice – Patis racemosa

Four-leaved milkweed – Asclepias quadrifolia

Rare and Uncommon Plants

Yellow oak – Quercus muehlenbergii

Wood lily – Lilium philadelphicum

Hitchcock’s sedge – Carex hitchcockiana

Forked chickweed – Paronychia canadensis

Allegheny crowfoot – Ranunculus allegheniensis

Early blue violet – Viola palmata

Deerberry – Vaccinium stamineum

Hound’s tongue – Cynoglossum boreale

Hairy honeysuckle – Lonicera hirsuta

Rue anemone – Thalictrum thalictroides

Back’s sedge – Carex backii

Rock muhlenbergia – Muhlenbergia sobolifera

Slender-flowered muhlenbergia – Muhlenbergia tenuiflora

Pignut hickory – Carya glabra

Early blue violet – Viola palmata

Rattlesnake-weed – Hieracium venosum

Downy arrowwood – Viburnum rafinesquianum

Kalm’s brome grass – Bromus kalmii

Fragrant sumac – Rhus aromatica

Longleaf bluet – Houstonia longifolia

Associated Animals

Eastern red-backed salamander – Plethodon cinereus 

Eastern gray squirrel – Sciurus carolinensis 

Southern flying squirrel – Glaucomys volans

Eastern chipmunk – Tamias striatus 

White-breasted nuthatch – Sitta carolinensis 

Scarlet tanager – Piranga olivacea 

Eastern wood pewee – Contopus virens

Red-eyed vireo – Vireo olivaceus 

Ovenbird – Seiurus aurocapilla

Baltimore oriole – Icterus galbula 

Wood thrush – Hylocichla mustelina 

Wild turkey – Meleagris gallopavo 

Hairy woodpecker – Picoides villosus 

Rose-breasted grosbeak – Pheucticus ludovicianus 

Tufted titmouse – Baeolophus bicolor 

Rare and Uncommon Animals

Eastern ratsnake – Pantherophis alleghaniensis

Common five-lined skink – Plestiodon fasciatus

Timber rattlesnake – Crotalus horridus 

Indiana bat – Myotis sodalist

Northern long-eared bat – Myotis septentrionalis 

Hickory hairstreak – Satyrium caryaevorus

Places to Visit

Shaw Mountain Natural Area, Benson, The Nature Conservancy (TNC).



Helen W. Buckner Memorial Preserve at Bald Mountain, West Haven, TNC



Mount Philo State Park, Charlotte, Vermont Department of Forests, Parks, and Recreation



Little Ascutney Wildlife Management Area, Weathersfield, Vermont Fish and Wildlife Department